1996-1998 Master course, Department of Pharmacology, Hokkaido University.
1998 Master of Pharmacology in Hokkaido University.
1998-2001 Doctor course, Department of Bioscience, Nara Institute of Science and Technology.
2001 Ph. D. in Nara Institute of Science and Technology.
2001-2003 Post-doctoral fellow in the Research Institute for Microbial Diseases, Osaka University
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