1993 PhD. candidate at School of Medicine, Kyoto University
1995 Honorary fellow/Research intern at Department of Biomolecular Chemistry, University of Wisconsin-Madison
1998 Postdoctoral fellow at Kitasato Institute
2000 Research fellow at Institute of scientific and industrial research, Osaka University
2001 Teaching assistant at Research Institute for Microbial Disease, Osaka University
2003 Research associate at Research Institute for Microbial Disease, Osaka University
2006 Research associate of COE at Research Institute for Microbial Disease, Osaka University
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